As of 2014, the "most comprehensive phylogenomic analysis of syndermatan relationships" to date was based on transcriptome data from all four groups, and provided "strong support" for the hypothesis illustrated in the bottom left of the figure, in which Seisonidea and Acanthocephala are sister taxa.The study further indicated that the sister group to this taxon is Bdelloidea, whereas Monogononta is the outgroup to all three.
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The main characteristics that distinguish bdelloids from related groups of rotifers are exclusively parthenogenetic reproduction and the ability to survive in dry, harsh environments by entering a state of desiccation-induced dormancy (anhydrobiosis) at any life stage. Modern molecular phylogenetic studies demonstrate that this classic understanding of 'Rotifera' is incomplete (paraphyletic), because it omits a fourth clade of closely related organisms: the Acanthocephala, or thorny-headed worms.
support a refined version of the scenario in the bottom left, with Bdelloidea as a sister group to Seisonidea Acanthocephala, and Monogononta as an outgroup. Originally classified as a separate phylum, molecular and morphological evidence accumulated between 19 to indicate that Acanthocephala forms a monophyletic group with Bdelloidea, Monogononta and Seisonidea.
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